Jane Goodall, one of the world’s most revered conservationists, who earned scientific stature and global celebrity by chronicling the distinctive behavior of wild chimpanzees in East Africa — primates that made and used tools, ate meat, held rain dances and engaged in organized warfare — died on Wednesday in Los Angeles. She was 91.

Her death, while on a speaking tour, was confirmed by the Jane Goodall Institute, whose U.S. headquarters are in Washington, D.C.

The British-born Dr. Goodall was 29 in the summer of 1963 when the National Geographic Society, which was financially supporting her field studies in the Gombe Stream Chimpanzee Reserve in what is now Tanzania, published her 7,500-word, 37-page account of the lives of Flo, David Greybeard, Fifi and other members of the troop of primates she had observed.

The article, with photographs by Hugo van Lawick, a Dutch wildlife photographer whom she later married, also described her struggles to overcome disease, predators and frustration as she tried to get close to the chimps, working from a primitive research station along the eastern shore of Lake Tanganyika.